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I believe keyframes always record the scale, position, and rotation of an object if you touch the object at all. It's unclear exactly what the relationship is with using FK (L2 or using the moves shouldn't make any difference to this), and the procedural run animations. It could be that they work simply because it's a rotation blending with a rotation, or perhaps it's relative to the connector or something? Hard to say...
Did you scale the keyframe with the puppet? I doubt that's it, but might be worth checking.